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It turns out the resolution was the [Network Acceleration](https://docs.gl-inet.com/router/en/4/interface_guide/network_acceleration/) option. It is enabled and set to Hardware Acceleration by default. Setting it to Software Acceleration or turning it off entirely resolves the issue. I currently just have it completely disabled.

I hadn't tried changing this option earlier, because it seemed counter-productive. In my case, it doesn't seem to cause any significant performance decrease for the WiFi connected devices and also does not seem to cause significant CPU load on the router itself. A benefit of *not* using Network Acceleration is that the speeds reported in the UI are accurate.

I'm not sure *why* Hardware Acceleration causes this issue. The [documentation mentions](https://docs.gl-inet.com/router/en/4/interface_guide/hardware_acceleration/) that it can interact poorly with Smart Queue Management (SQM). I had set up SQM on the device directly through LuCI, but disabling it and changing queuing discipline settings on the desktop didn't seem to resolve the issue. I don't know if merely *installing* the SQM package on the router caused the issue. Again, the devices connected via WiFi did see the latency differences of enabling SQM without having their performance destroyed.